Models whose bases are touching are engaged in close combat. This can only happen once a warrior has charged his enemy, as models are otherwise not allowed to move into contact.
All close quarter fighting is worked out in the Close Combat phase. Regardless of whose turn it is, all models in close combat will fight. A warrior can fight against enemies to his side, front, or rear. In reality the fighters are constantly moving, dodging, and weaving as they struggle to kill their adversaries. Models fighting in close combat do not shoot in the Shooting phase. They are far too busy fighting for their lives. Any very close range shots they are able to make using pistols are treated like close combat weapon attacks (see the Weapons & Amour section).